Biology ECOSYSTEM

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ECOSYSTEM

Productivity

A constant input of solar energy is the basic requirement for any ecosystem to function and sustain. Primary production is defined as the amount of biomass or organic matter produced per unit area over a time period by plants during photosynthesis. It is expressed in terms of weight (g–2) orenergy (kcal m–2). The rate of biomass production is called productivity.
It is expressed in terms of g–2 yr–1 or (kcal m–2) yr–1 to compare the productivity of different ecosystems. It can be divided into gross primary productivity (GPP) and net primary productivity (NPP). Gross primary productivity of an ecosystem is the rate of production of organic matter during photosynthesis. A considerable amount of GPP is utilised by plants in respiration. Gross primary productivity minus respiration losses (R), is the net primary productivity (NPP).
GPP – R = NPP
Net primary productivity is the available biomass for the consumption to heterotrophs (herbiviores and decomposers). Secondary productivity is defined as the rate of formation of new organic matter by consumers.
Primary productivity depends on the plant species inhabiting a particular area. It also depends on a variety of environmental factors,
availability of nutrients and photosynthetic capacity of plants. Therefore, it varies in different types of ecosystems. The annual net primary
productivity of the whole biosphere is approximately 170 billion tons (dry weight) of organic matter. Of this, despite occupying about 70 per cent of the surface, the productivity of the oceans are only 55 billion tons. Rest of course, is on land.

A constant input of solar energy is the basic requirement for any ecosystem to function and sustain. Primary production is defined as the amount of biomass or organic matter produced per unit area over a time period by plants during photosynthesis. It is expressed in terms of weight (g–2) orenergy (kcal m–2). The rate of biomass production is called productivity.
It is expressed in terms of g–2 yr–1 or (kcal m–2) yr–1 to compare the productivity of different ecosystems. It can be divided into gross primary productivity (GPP) and net primary productivity (NPP). Gross primary productivity of an ecosystem is the rate of production of organic matter during photosynthesis. A considerable amount of GPP is utilised by plants in respiration. Gross primary productivity minus respiration losses (R), is the net primary productivity (NPP).
GPP – R = NPP
Net primary productivity is the available biomass for the consumption to heterotrophs (herbiviores and decomposers). Secondary productivity is defined as the rate of formation of new organic matter by consumers.
Primary productivity depends on the plant species inhabiting a particular area. It also depends on a variety of environmental factors,
availability of nutrients and photosynthetic capacity of plants. Therefore, it varies in different types of ecosystems. The annual net primary
productivity of the whole biosphere is approximately 170 billion tons (dry weight) of organic matter. Of this, despite occupying about 70 per cent of the surface, the productivity of the oceans are only 55 billion tons. Rest of course, is on land.

DECOMPOSITION

You may have heard of the earthworm being referred to as the farmer’s ‘friend’. This is so because they help in the breakdown of complex organic matter as well as in loosening of the soil. Similarly, decomposers break down complex organic matter into inorganic substances like carbon dioxide, water and nutrients and the process is called decomposition.
Dead plant remains such as leaves, bark, flowers and dead remains of animals, including fecal matter, constitute detritus, which is the raw material for decomposition. The important steps in the process of decomposition are fragmentation, leaching, catabolism, humification and mineralisation.
Detritivores (e.g., earthworm) break down detritus into smaller particles. This process is called fragmentation. By the process of leaching, watersoluble inorganic nutrients go down into the soil horizon and get precipitated as unavailable salts. Bacterial and fungal enzymes degrade detritus into simpler inorganic substances. This process is called as catabolism.
It is important to note that all the above steps in decomposition operate simultaneously on the detritus (Figure 14.1). Humification and
mineralisation occur during decomposition in the soil. Humification leads to accumulation of a dark coloured amorphous substance called humus that is highly resistant to microbial action and undergoes decomposition at an extremely slow rate. Being colloidal in nature it serves as a reservoir of nutrients. The humus is further degraded by some microbes and release of inorganic nutrients occur by the process known as mineralisation.
Decomposition is largely an oxygen-requiring process. The rate of decomposition is controlled by chemical composition of detritus and
climatic factors. In a particular climatic condition, decomposition rate is slower if detritus is rich in lignin and chitin, and quicker, if detritus is
rich in nitrogen and water-soluble substances like sugars. Temperature and soil moisture are the most important climatic factors that regulate decomposition through their effects on the activities of soil microbes. Warm and moist environment favour decomposition whereas low temperature and anaerobiosis inhibit decomposition resulting in build up of organic materials.

You may have heard of the earthworm being referred to as the farmer’s ‘friend’. This is so because they help in the breakdown of complex organic matter as well as in loosening of the soil. Similarly, decomposers break down complex organic matter into inorganic substances like carbon dioxide, water and nutrients and the process is called decomposition.
Dead plant remains such as leaves, bark, flowers and dead remains of animals, including fecal matter, constitute detritus, which is the raw material for decomposition. The important steps in the process of decomposition are fragmentation, leaching, catabolism, humification and mineralisation.
Detritivores (e.g., earthworm) break down detritus into smaller particles. This process is called fragmentation. By the process of leaching, watersoluble inorganic nutrients go down into the soil horizon and get precipitated as unavailable salts. Bacterial and fungal enzymes degrade detritus into simpler inorganic substances. This process is called as catabolism.
It is important to note that all the above steps in decomposition operate simultaneously on the detritus (Figure 14.1). Humification and
mineralisation occur during decomposition in the soil. Humification leads to accumulation of a dark coloured amorphous substance called humus that is highly resistant to microbial action and undergoes decomposition at an extremely slow rate. Being colloidal in nature it serves as a reservoir of nutrients. The humus is further degraded by some microbes and release of inorganic nutrients occur by the process known as mineralisation.
Decomposition is largely an oxygen-requiring process. The rate of decomposition is controlled by chemical composition of detritus and
climatic factors. In a particular climatic condition, decomposition rate is slower if detritus is rich in lignin and chitin, and quicker, if detritus is
rich in nitrogen and water-soluble substances like sugars. Temperature and soil moisture are the most important climatic factors that regulate decomposition through their effects on the activities of soil microbes. Warm and moist environment favour decomposition whereas low temperature and anaerobiosis inhibit decomposition resulting in build up of organic materials.

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